Flora of the Hawaiian Islands
Dicotyledon Literature for Melicope clusiifolia
Wagner et al., 1990, 1999.
   Rutaceae -- The Rue Family Bibliography
      Melicope clusiifolia

Common name(s): alani, alani kuahiwi, kolokolo mokihana, kuaemoa (Kaua`i)
General Information
DistributionIn the Hawaiian Islands, endemic to Kaua`i, O`ahu, Moloka`i, Lana`i, Maui, Hawai`i.

Click here for detailed USGS map by Jonathan Price
Shrubs or trees 2-10 m tall, new growth glabrous to minutely scurfy and hirsute or densely yellowish brown or golden pilose to shaggy pilose.
Leaves with a weak anise odor, in whorls of (2-)4(-8) per node, subcoriaceous to coriaceous, often brittle, variable in shape, obovate, oblong-spatulate, or elliptic-obovate, rarely oblong-elliptic, elliptic-ovate, or elliptic, (2.5-)5-30 cm long, (1.5-)2.5-6(-7.5) cm wide, primary lateral veins usually 14-20 pairs, connected by a weakly arched vein ca. 1-4 mm from margin, primary lateral veins and submarginal vein only slightly more prominent than higher order venation, which forms an evident reticulate pattern, upper surface usually glossy, glabrous or midrib sometimes short-hirtellous, lower surface usually paler, glabrate to puberulent or short-villous along midrib, sometimes more evenly over entire surface, margins sometimes recurved or revolute, apex rounded to subtruncate and usually emarginate, sometimes obtuse, base rounded, cuneate, or attenuate, petioles 0-40 mm long.
Flowers perfect or functionally unisexual (and then plants dioecious), in glabrous, fasciculate cymes, initiated among leaves, maturing usually on naked stems below leaves, peduncles 0-4 mm long, pedicels 1-10 mm long, bracteoles minute, deltate; staminate flowers with calyx ovate, 3-4 mm long, glabrous or minutely ciliate, petals deltate-ovate, 4-8 mm long, glabrous, apex acute, ovary rudimentary, glabrous; pistillate flowers with sepals ovate, 2-3 mm long, glabrous or minutely ciliate, petals deltate, 3-6 mm long, glabrous, ovary glabrous, style ca. 2 mm long, stigma lobes thick, blunt.
Capsules 8-18 mm wide, carpels connate 1/8-7/8 their length, 4-9 mm long, exocarp usually strongly wrinkled when dry, glabrous, endocarp glabrous.
Seeds 1-2 per carpel, 2.5-5(-6) mm long.