Sicyos microcarpus H. Mann var. laysanensis (H. St. John) H. St. John, Phytologia 63: 190. 1987. Flora of the Hawaiian Islands --Basionym
Type Information
Sicyos laysanensis H. St. John , Pacific Sci. 24: 440 1970.
Laysan. May 1911. W. A. Bryan A.
Holotype [BISH]. Isotype(s) [BISH!].

Note: Two other specimens, collected April 1903, were annotated as "isotype" by St. John but are not isotypes.

=Sicyos pachycarpus
*Verification: Wagner, W.L. & R.K. Shannon. 1999. Novon 9:441-447.
---------------------


Note: Published simultaneously with S. microcarpus subsp. laysanensis.

*Verification: Wagner, W.L. & R.K. Shannon. 1999. Novon 9:441-447.

Sicyos microcarpus H. Mann var. meeboldii H. St. John, Phytologia 63: 191. 1987.
Type information:
Holotype [BISH]. G.L. Webster and R.L. Wilbur 1837. 23 June 1948. Hawai`i: North Kona, Kaupulehu, 3 mi. E of Huehue.2000ft. Isotype(s) [TEX].

Note: St. John erroneously published the collector number as Webster 1537, which is a specimen of Phyllostegia grandiflora collected on O`ahu.

=Sicyos pachycarpus
Note: St. John erroneously published the collector number as Webster 1537, which is a specimen of Phyllostegia grandiflora collected on O`ahu.

*Verification: Wagner, W.L. & R.K. Shannon. 1999. Novon 9:441-447.

Sicyos microcarpus H. Mann var. obovoideis H. St. John, Phytologia 63: 191. 1987.
Type information:
Holotype [BISH]. H. F. Clay s.n.. s.d.. Maui: s.l.. Isotype(s) [BISH!, PTBG!, US!].

Note: Although published by St. John as collected by Clay, the label clearly states: "Sent from Maui to H. F. Clay; given by him to Bishop Museum May 16, 1954." Published as "obovoideis," but St. John's handwriting on the specimen label is "obovoideus."

=Sicyos pachycarpus
Note: Although published by St. John as collected by Clay, the label clearly states: "Sent from Maui to H. F. Clay; given by him to Bishop Museum May 16, 1954." Published as "obovoideis," but St. John's handwriting on the specimen label is "obovoideus."

*Verification: Wagner, W.L. & R.K. Shannon. 1999. Novon 9:441-447.

Sicyos microcarpus H. Mann var. oniniensis H. St. John, Phytologia 63: 190. 1987.
Type information:
Holotype [BISH]. L. E. Bishop 1690. 21 January 1973. Moloka`i: Onini Gulch bottom.2200ft.
=Sicyos pachycarpus

*Verification: Wagner, W.L. & R.K. Shannon. 1999. Novon 9:441-447.

Sicyos microcarpus H. Mann var. paneensis H. St. John, Phytologia 63: 188. 1987.
Type information:
Holotype [NY]. O. Degener 18035. 12 April 1933. O`ahu: Kamokuiki Valley, between Pu`u Iki and Pu`u Kamaohanui. Isotype(s) [NY! [2]].
=Sicyos pachycarpus

*Verification: Wagner, W.L. & R.K. Shannon. 1999. Novon 9:441-447.

Sicyos microcarpus H. Mann var. pohakeaensis H. St. John, Phytologia 63: 188. 1987.
Type information:
Holotype [NY]. O. Degener 18400. 11 July 1927. Maui: West Maui, Pohakea Gulch, arid valley floor.
=Sicyos pachycarpus

*Verification: Wagner, W.L. & R.K. Shannon. 1999. Novon 9:441-447.

Sicyos microcarpus H. Mann var. trilobata H. St. John, Phytologia 63: 190. 1987.
Type information:
Holotype [BISH]. W. C. Gagné 676. 15 May 1973. Moloka`i: Makolelau Gulch.760m.
=Sicyos pachycarpus

*Verification: Wagner, W.L. & R.K. Shannon. 1999. Novon 9:441-447.

Sicyos microcarpus H. Mann var. woolfordii H. St. John, Phytologia 63: 189. 1987.
Type information:
Holotype [BISH]. H. A. Woolford s.n.. 11 March 1957. O`ahu: `Ewa coral flats. Isotype(s) [BISH!].
=Sicyos pachycarpus

*Verification: Wagner, W.L. & R.K. Shannon. 1999. Novon 9:441-447.

Sicyos microcarpus H. Mann , Proc. Amer. Acad. Arts 7: 167. 1867.
Type information:
Holotype [A]. J. Rémy 541. 1851- 1855. O`ahu. Isotype(s) [BR!, P!].
=Sicyos pachycarpus

*Verification: Wagner, W.L. & R.K. Shannon. 1999. Novon 9:441-447.

Sicyos microcarpus H. Mann subsp. kanepuuensis H. St. John, Phytologia 63: 189. 1987.
Type information:
Holotype [BISH]. G.C. Munro 10. 28 May 1927. Lana`i: Kanepu`u. Isotype(s) [BISH!].
=Sicyos pachycarpus

*Verification: Wagner, W.L. & R.K. Shannon. 1999. Novon 9:441-447.

Sicyos microcarpus H. Mann subsp. lanaiensis H. St. John, Phytologia 63: 189. 1987.
Type information:
Holotype [BISH]. G.C. Munro 535. 24 March 1916. Lana`i: Kolulu. Isotype(s) [BISH! [3]].

Note: Two of the isotypes have additional collection data, possibly written by Forbes, including a date of "3/29/16." This appears to be a misinterpretation of the original handwritten label on the holotype. Published simultaneously with S. microcarpus var. lanaiensis.

=Sicyos pachycarpus
Note: Two of the isotypes have additional collection data, possibly written by Forbes, including a date of "3/29/16." This appears to be a misinterpretation of the original handwritten label on the holotype. Published simultaneously with S. microcarpus var. lanaiensis.

*Verification: Wagner, W.L. & R.K. Shannon. 1999. Novon 9:441-447.

Sicyos microcarpus H. Mann subsp. laysanensis (H. St. John) H. St. John, Phytologia 63: 190. 1987. Flora of the Hawaiian Islands --Basionym
Type Information
Sicyos laysanensis H. St. John , Pacific Sci. 24: 440 1970.
Laysan. May 1911. W. A. Bryan A.
Holotype [BISH]. Isotype(s) [BISH!].

Note: Two other specimens, collected April 1903, were annotated as "isotype" by St. John but are not isotypes.

=Sicyos pachycarpus
*Verification: Wagner, W.L. & R.K. Shannon. 1999. Novon 9:441-447.
---------------------


Note: Published simultaneously with S. microcarpus var. laysanensis.

*Verification: Wagner, W.L. & R.K. Shannon. 1999. Novon 9:441-447.

Sicyos microcarpus H. Mann subsp. puberulus H. St. John, Phytologia 63: 191. 1987.
Type information:
Holotype [US]. O. Degener 18391. 10 April 1932. O`ahu: Wai`anae, arid roadside near town. Isotype(s) [NY! [2]].
=Sicyos pachycarpus

*Verification: Wagner, W.L. & R.K. Shannon. 1999. Novon 9:441-447.

Sicyos microcarpus H. Mann var. ewaensis H. St. John, Phytologia 63: 190. 1987.
Type information:
Holotype [BISH]. K. M. Nagata and B. Kimura 1656. 21 June 1978. O`ahu: west side of coral plains at Ewa, near Barber's Point, ocean side of Malakole Rd. near Camp Malakole Military Res..10ft. Isotype(s) [BISH!].
=Sicyos pachycarpus

*Verification: Wagner, W.L. & R.K. Shannon. 1999. Novon 9:441-447.

Sicyos microcarpus H. Mann var. forbesii H. St. John, Phytologia 63: 188. 1987.
Type information:
Holotype [BISH]. C.N. Forbes 11.M. May 1910. Maui: West Maui, Maunaho`oma, [in gulch].
=Sicyos pachycarpus

*Verification: Wagner, W.L. & R.K. Shannon. 1999. Novon 9:441-447.

Sicyos microcarpus H. Mann var. haleakalaensis H. St. John, Phytologia 63: 191. 1987.
Type information:
Holotype [BISH]. C.N. Forbes 1851.M. 5 March 1920. Maui: [East Maui], Wai`opai, [gulch on rt. hand side of camp].

Note: Date from Forbes's field book. Holotype missing at least since October 1987.

=Sicyos pachycarpus
Note: Date from Forbes's field book. Holotype missing at least since October 1987.

*Verification: Wagner, W.L. & R.K. Shannon. 1999. Novon 9:441-447.

Sicyos microcarpus H. Mann var. kalaupapaensis H. St. John, Phytologia 63: 188. 1987.
Type information:
Holotype [BISH]. J. F. Rock 14061. 24 May 1918. Moloka`i: Kalaupapa beach. Isotype(s) [BISH! [2]].
=Sicyos pachycarpus

*Verification: Wagner, W.L. & R.K. Shannon. 1999. Novon 9:441-447.

Sicyos microcarpus H. Mann var. kamoaensis H. St. John, Phytologia 63: 189. 1987.
Type information:
Holotype [BISH]. G.C. Munro 418. 16 May 1929. Lana`i: Kamoa.1740ft. Isotype(s) [BISH! [3]].
=Sicyos pachycarpus

*Verification: Wagner, W.L. & R.K. Shannon. 1999. Novon 9:441-447.

Sicyos microcarpus H. Mann var. kauensis H. St. John, Phytologia 63: 191. 1987.
Type information:
Holotype [BISH]. J. F. Rock s.n.. April 1911. Hawai`i: Ka`u, on lava.
=Sicyos pachycarpus

*Verification: Wagner, W.L. & R.K. Shannon. 1999. Novon 9:441-447.

Sicyos microcarpus H. Mann var. kealiaensis H. St. John, Phytologia 63: 189. 1987. Flora of the Hawaiian Islands --Basionym
Type Information

Note: Not validly published. W. C. Gagné 671 (BISH! [2]) was cited as the type collection for this name and in the same publication it was cited again as the type collection for S. erostratus var. kealiaensis, therefore neither name is validly published under ICBN Art. 34.2 (Greuter et. al., 1994). Telford placed S. microcarpus var. kealiaensis in the synonomy of S. pachycarpus.

*Verification: Wagner, W.L. & R.K. Shannon. 1999. Novon 9:441-447.

Sicyos microcarpus H. Mann var. kilaueaensis H. St. John, Phytologia 63: 188. 1987.
Type information:
Holotype [BISH]. G.O. Fagerlund and A.M. Mitchell 544. 10 May 1943. Hawai`i: Apua, Hawai`i [Volcanoes] National Park, on slope of pali.2000ft.
=Sicyos pachycarpus

*Verification: Wagner, W.L. & R.K. Shannon. 1999. Novon 9:441-447.

Sicyos microcarpus H. Mann var. koeleensis H. St. John, Phytologia 63: 190. 1987.
Type information:
Holotype [BISH]. C.N. Forbes 95.L. 9 June 1913. Lana`i: Mts. near Ko`ele. Isotype(s) [BISH! [4]].

Note: Date from Forbes's field books.

=Sicyos pachycarpus
Note: Date from Forbes's field books.

*Verification: Wagner, W.L. & R.K. Shannon. 1999. Novon 9:441-447.

Sicyos microcarpus H. Mann var. koloaensis H. St. John, Phytologia 63: 191. 1987.
Type information:
Holotype [BISH]. C.H. Lamoureux 2346. 13 April 1963. Kaua`i: Koloa Dist., ahupua`a of Koloa, 1 mi. W of Poipu Beach Park.10ft. Isotype(s) [BISH! [3]].
=Sicyos pachycarpus

*Verification: Wagner, W.L. & R.K. Shannon. 1999. Novon 9:441-447.

Sicyos microcarpus H. Mann var. lanaiensis H. St. John, Phytologia 63: 189. 1987.
Type information:
Holotype [BISH]. G.C. Munro 535. 24 March 1916. Lana`i: Kolulu. Isotype(s) [BISH! [3]].

Note: Two of the isotypes have additional collection data, possibly written by Forbes, including a date of "3/29/16." This appears to be a misinterpretation of the original handwritten label on the holotype. Published simultaneously with S. microcarpus subsp. lanaiensis.

=Sicyos pachycarpus
Note: Two of the isotypes have additional collection data, possibly written by Forbes, including a date of "3/29/16." This appears to be a misinterpretation of the original handwritten label on the holotype. Published simultaneously with S. microcarpus subsp. lanaiensis.

*Verification: Wagner, W.L. & R.K. Shannon. 1999. Novon 9:441-447.