Gouldia terminalis (Hook. & Arn.) Hillebr. , Fl. Hawaiian Isl. 169. 1888. Flora of the Hawaiian Islands --Basionym
Type Information

*Verification: D.M. Mix.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. acuminata Fosberg, Bernice P. Bishop Mus. Bull. 147: 45. 1937.
Type information:
Holotype [BISH]. C.N. Forbes 190.M. 28-9 June-July 1910. Maui: East Maui, N and NW slopes of Haleakala, above Olinda, woods near Ukulele, wet forests.

Note: Date from Forbes's field book.

=Kadua affinis
Note: Date from Forbes's field book.

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. acuta Fosberg, Bernice P. Bishop Mus. Bull. 147: 55. 1937.
Type information:
Holotype [BISH]. F.R. Fosberg 10125. 30 August 1933. Hawai`i: Kilauea, Kipuka Puaulu (Bird Park), dry open forest.1230m.
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. albicaulis Fosberg, Bernice P. Bishop Mus. Bull. 147: 32. 1937.
Type information:
Holotype [BISH]. C.N. Forbes 98.K. 29 July 1909. Kaua`i: ridge west of Waiali Valley.
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. cuneata Fosberg, Bernice P. Bishop Mus. Bull. 147: 35. 1937.
Type information:
Holotype [BISH]. H. St. John and F.R. Fosberg 13769. 27 December 1933. Kaua`i: Waimea, Milolii Ridge, wet forest.3100ft. Isotype(s) [BISH!].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. euantiqua Fosberg, Bernice P. Bishop Mus. Bull. 147: 54. 1937. Flora of the Hawaiian Islands --Basionym
Type Information

Note: Not validly published under the provisions of Art. 24.3, ICBN 1994.

*Verification: R.K. Shannon, W.L. Wagner, D.H. Nicolson.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. euarborescens Fosberg, Bernice P. Bishop Mus. Bull. 147: 31. 1937. Flora of the Hawaiian Islands --Basionym
Type Information

Note: Not validly published under the provisions of Art. 24.3, ICBN 1994.

*Verification: R.K. Shannon, W.L. Wagner, D.H. Nicolson.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. eucordata Fosberg, Bernice P. Bishop Mus. Bull. 147: 44. 1937. Flora of the Hawaiian Islands --Basionym
Type Information

Note: Not validly published under the provisions of Art. 24.3, ICBN 1994.

*Verification: R.K. Shannon, W.L. Wagner, D.H. Nicolson.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. euelongata Fosberg, Bernice P. Bishop Mus. Bull. 147: 32. 1937. Flora of the Hawaiian Islands --Basionym
Type Information

Note: Not validly published under the provisions of Art. 24.3, ICBN 1994.

*Verification: R.K. Shannon, W.L. Wagner, D.H. Nicolson.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. euglabra Fosberg, Bernice P. Bishop Mus. Bull. 147: 36. 1937. Flora of the Hawaiian Islands --Basionym
Type Information

Note: Not validly published under the provisions of Art. 24.3, ICBN 1994.

*Verification: R.K. Shannon, W.L. Wagner, D.H. Nicolson.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. eukaala Fosberg, Bernice P. Bishop Mus. Bull. 147: 49. 1937. Flora of the Hawaiian Islands --Basionym
Type Information

Note: Not validly published under the provisions of Art. 24.3, ICBN 1994.

*Verification: R.K. Shannon, W.L. Wagner, D.H. Nicolson.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. eukapuaensis Fosberg, Bernice P. Bishop Mus. Bull. 147: 39. 1937. Flora of the Hawaiian Islands --Basionym
Type Information

Note: Not validly published under the provisions of Art. 24.3, ICBN 1994.

*Verification: R.K. Shannon, W.L. Wagner, D.H. Nicolson.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. eukonaensis Fosberg, Bernice P. Bishop Mus. Bull. 147: 38. 1937. Flora of the Hawaiian Islands --Basionym
Type Information

Note: Not validly published under the provisions of Art. 24.3, ICBN 1994.

*Verification: R.K. Shannon, W.L. Wagner, D.H. Nicolson.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. eumacrocarpa Fosberg, Bernice P. Bishop Mus. Bull. 147: 34. 1937. Flora of the Hawaiian Islands --Basionym
Type Information

Note: Not validly published under the provisions of Art. 24.3, ICBN 1994.

*Verification: R.K. Shannon, W.L. Wagner, D.H. Nicolson.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. euovata Fosberg, Bernice P. Bishop Mus. Bull. 147: 50. 1937. Flora of the Hawaiian Islands --Basionym
Type Information

Note: Not validly published under the provisions of Art. 24.3, ICBN 1994.

*Verification: R.K. Shannon, W.L. Wagner, D.H. Nicolson.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. euparvifolia Fosberg, Bernice P. Bishop Mus. Bull. 147: 56. 1937. Flora of the Hawaiian Islands --Basionym
Type Information

Note: Not validly published under the provisions of Art. 24.3, ICBN 1994.

*Verification: R.K. Shannon, W.L. Wagner, D.H. Nicolson.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. eustipulacea Fosberg, Bernice P. Bishop Mus. Bull. 147: 43. 1937. Flora of the Hawaiian Islands --Basionym
Type Information

Note: Not validly published under the provisions of Art. 24.3, ICBN 1994.

*Verification: R.K. Shannon, W.L. Wagner, D.H. Nicolson.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. gracilis Fosberg, Bernice P. Bishop Mus. Bull. 147: 29. 1937.
Type information:
Holotype [BISH]. E.P. Hume 345. 28 November 1931. O`ahu: Ko`olau Range, Manoa, Manoa Cliff trail, shaded gulch.1800ft.

Note: Collection month published as December, but label says "11/28/31."

=Kadua affinis
Note: Collection month published as December, but label says "11/28/31."

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. heterophylla Fosberg, Bernice P. Bishop Mus. Bull. 147: 52. 1937.
Type information:
Holotype [BISH]. H. St. John and F.R. Fosberg 13091. 29 December 1932. Moloka`i: trail on ridge E of Mapulehu Valley, wet forest.2100ft. Isotype(s) [BISH!].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. hirtellicostata Fosberg, Bernice P. Bishop Mus. Bull. 147: 33. 1937.
Type information:
Holotype [BISH]. H. St. John and F.R. Fosberg 13550. 24 December 1933. Kaua`i: Kalaheo, slopes of Kahili, above Kahili Bog.2200ft. Isotype(s) [BISH!].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. hirtellifolia Fosberg, Bernice P. Bishop Mus. Bull. 147: 55. 1937.
Type information:
Holotype [BISH]. C.N. Forbes 327.H. 15 July 1911. Hawai`i: Kona, Papaloa. Isotype(s) [BISH! [2]].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. impressa Fosberg, Bull. Torrey Bot. Club 70: 391. 1943.
Type information:
Holotype [NY]. O. Degener 12521. 5 August 1939. Maui: [East Maui], Haleakala, Paliku, decadent forest. Isotype(s) [MO!, NY [mfc!], US!].

Note: Published as a forma of var. parvula, but the other formas he compares it to are formas of var. parvifolia, and he annotated the NY and US sheets as G. terminalis var. parvifolia f. impressa.

=Kadua affinis
Note: Published as a forma of var. parvula, but the other formas he compares it to are formas of var. parvifolia, and he annotated the NY and US sheets as G. terminalis var. parvifolia f. impressa.

*Verification: R.K. Shannon.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. kahili Fosberg, Bernice P. Bishop Mus. Bull. 147: 33. 1937.
Type information:
Holotype [BISH]. H. St. John and F.R. Fosberg 13532. 24 December 1933. Kaua`i: Kalaheo, slopes of Kahili, above Kahili bog, wet forest.2400ft. Isotype(s) [BISH!].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. kalaupapa Fosberg, Bernice P. Bishop Mus. Bull. 147: 51. 1937.
Type information:
Holotype [BISH]. C.N. Forbes 25.Mo. June 1912. Moloka`i: on cliffs above Kalaupapa, Settlement Trail. Isotype(s) [BISH! [3], US!].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. kauensis Fosberg, Bernice P. Bishop Mus. Bull. 147: 55. 1937.
Type information:
Holotype [BISH]. C.N. Forbes 384.H. 2 August 1911. Hawai`i: Ka`u Desert. Isotype(s) [BISH!, US!].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. kehena Fosberg, Bernice P. Bishop Mus. Bull. 147: 54. 1937.
Type information:
Holotype [BISH]. F.R. Fosberg 10195. 2 September 1933. Hawai`i: Kohala Mts., Kehena Ditch Trail, near Kehena.770m.
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. latifolia Fosberg, Bernice P. Bishop Mus. Bull. 147: 38. 1937.
Type information:
Holotype [BISH]. J. F. Rock 3866. 18 June 1909. Hawai`i: North Kona, Puuwaawaa.

Note: Apparently Rock's name Gouldia grandifolia was never published.

=Kadua affinis
Note: Apparently Rock's name Gouldia grandifolia was never published.

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. lydgatei Fosberg, Bernice P. Bishop Mus. Bull. 147: 53. 1937.
Type information:
Holotype [BISH]. W. Hillebrand and J.M. Lydgate 126. 1851- 1871. Maui: Hamakua.

Note: The sheet containing the holotype represents a mixed collection of three different specimens. The twig in the upper left is not a Rubiaceae; the twig in the upper right is apparently from a different locality; only the twig in the lower left is the holotype. The publication indicates that there is a photo of the type at BISH, but it could not be located (RKS, 8/95). There is a photo of a sheet that Fosberg labelled "isotype," but it cannot be an isotype because it is from O`ahu, and the type was collected on Maui.

=Kadua affinis
Note: The sheet containing the holotype represents a mixed collection of three different specimens. The twig in the upper left is not a Rubiaceae; the twig in the upper right is apparently from a different locality; only the twig in the lower left is the holotype. The publication indicates that there is a photo of the type at BISH, but it could not be located (RKS, 8/95). There is a photo of a sheet that Fosberg labelled "isotype," but it cannot be an isotype because it is from O`ahu, and the type was collected on Maui.

*Verification: W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. macrophylla Fosberg, Bernice P. Bishop Mus. Bull. 147: 32. 1937.
Type information:
Holotype [BISH]. H. St. John and F.R. Fosberg 13597. 24 December 1933. Kaua`i: Wahiawa, Wahiawa Valley, trail to Kahili bog, wet forest.1900ft. Isotype(s) [BISH!].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. makawaoensis Fosberg, Bernice P. Bishop Mus. Bull. 147: 50. 1937.
Type information:
Holotype [BISH]. J. F. Rock 8531. September 1910. Maui: E Maui, Haleakala, Makawao. Isotype(s) [BISH!].

Note: Apparently Rock's name Gouldia haleakalensis was never published.

=Kadua affinis
Note: Apparently Rock's name Gouldia haleakalensis was never published.

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. maunahui Fosberg, Bernice P. Bishop Mus. Bull. 147: 51. 1937.
Type information:
Holotype [BISH]. H. St. John , R.J. Baker and J.W. Coulter, F.R. Fosberg, 12284. 20 December 1932. Moloka`i: Kawela, ridge between Maunahui, Kaunakakai, and Hanalilolilo.3640ft. Isotype(s) [BISH!].

Note: Additional collector: T. G. Yuncker.

=Kadua affinis
Note: Additional collector: T. G. Yuncker.

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. membranacea Fosberg, Bernice P. Bishop Mus. Bull. 147: 53. 1937.
Type information:
Holotype [BISH]. O. Degener 9478. 16 June 1927. Maui: Olinda, along pipeline trail, in rain forest. Isotype(s) [BISH!, NY [mfc!]].

Note: On label of BISH isotype is written (in Degener's handwriting), "Dupl. to NY, Fosberg." Check at US for isotype.

=Kadua affinis
Note: On label of BISH isotype is written (in Degener's handwriting), "Dupl. to NY, Fosberg." Check at US for isotype.

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. molokaiensis Fosberg, Bernice P. Bishop Mus. Bull. 147: 45. 1937.
Type information:
Holotype [BISH]. H. St. John 12580. 25 December 1932. Moloka`i: Kawela, Waikolu Valley, ridge between Hanalilolilo and Pepeopae, in wet boggy forest.4000ft. Isotype(s) [BISH! [3]].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. nealae Fosberg, Bernice P. Bishop Mus. Bull. 147: 45. 1937.
Type information:
Holotype [BISH]. C.N. Forbes 2668.M. 2 July 1920. Maui: E of Ke`anae, ridge between West and East Wailuaiki.
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. oahuensis Fosberg, Brittonia 8: 176. 1956.
Type information:
Holotype [NY]. O. Degener and W. Hatheway 21172. 14 December 1950. O`ahu: Mokuleia, Keawapilau, forest.1800ft. Isotype(s) [BISH!, US!].

Note: Publication gives collection number as 21722, but BISH, NY, and US sheets have 21172 on the label.

=Kadua affinis
Note: Publication gives collection number as 21722, but BISH, NY, and US sheets have 21172 on the label.

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. oblonga Fosberg, Brittonia 8: 176. 1956.
Type information:
Holotype [NY]. O. Degener and T. Murashige 20337. 13 August 1949. Hawai`i: Kona, Capt. Cook, S of Papaloa.5000ft. Isotype(s) [BISH! [2], US!].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. obovata Fosberg, Bull. Torrey Bot. Club 70: 391. 1943.
Type information:
Holotype [NA]. F.R. Fosberg 12472. 30 November 1935. Lana'i: between Maunalaei and Hauola drainages, ridge below Puu Aalii, rather dry forest.800m. Isotype(s) [BISH!, US!].

Note: Hauola spelled "Hauula" in publication.

=Kadua affinis
Note: Hauola spelled "Hauula" in publication.

*Verification: R.K. Shannon.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. ovata Fosberg, Brittonia 8: 176. 1956. Flora of the Hawaiian Islands --Basionym
Type Information

Note: Invalidly pulished according to provisions of ICBN Art. 33.2 (1994).

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. parvithyrsa Fosberg, Bernice P. Bishop Mus. Bull. 147: 37. 1937.
Type information:
Holotype [BISH]. U. Faurie 356. June 1909. Hawai`i: central part of Hamakua Coast, Kalua.
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. petiolata Fosberg, Bernice P. Bishop Mus. Bull. 147: 53. 1937.
Type information:
Holotype [BISH]. G.C. Munro 370. 5 June 1916. Moloka`i: Kalamaula.
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. pittosporoides Fosberg, Bernice P. Bishop Mus. Bull. 147: 39. 1937.
Type information:
Holotype [BISH]. J. F. Rock 10031. 28 January 1912. Hawai`i: Ka`u, Manuka.

Note: Only part of this collection is type for this entity; the other part was used as the type of Gouldia terminalis var. konaensis Fosb. Rock's name Gouldia volcanica was apparently never published.

=Kadua affinis
Note: Only part of this collection is type for this entity; the other part was used as the type of Gouldia terminalis var. konaensis Fosb. Rock's name Gouldia volcanica was apparently never published.

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. punaula Fosberg, Bernice P. Bishop Mus. Bull. 147: 53. 1937.
Type information:
Holotype [BISH]. H. St. John and F.R. Fosberg 12797. 28 December 1932. Moloka`i: ridge W of Punaula Valley, wooded ridge.1900ft. Isotype(s) [BISH! [3], US!].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. rigidifolia Fosberg, Bernice P. Bishop Mus. Bull. 147: 39. 1937.
Type information:
Holotype [BISH]. F.R. Fosberg 10093. 29 August 1933. Hawai`i: Kilauea, floor of Napau Crater, edge of new lava flow.750m. Isotype(s) [BISH!].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. rigidifolioides Fosberg, Brittonia 8: 174. 1956.
Type information:
Holotype [US]. O. Degener 21768. 3 February 1952. Hawai`i: near Kulani Prison Road, dense forest.4000ft.
=Kadua affinis

*Verification: R.K. Shannon.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. robusta Fosberg, Bernice P. Bishop Mus. Bull. 147: 29. 1937.
Type information:
Holotype [BISH]. C.N. Forbes s.n.. 10 December 1908. O`ahu: Koolau Mts., Waiolani ridge--Lanihuli Trail. Isotype(s) [BISH!].

Note: Although the two sheets have slightly different locality data we assume they are the same collection, based on the itinerary and list of collections given for that date in Forbes's field book.

=Kadua affinis
Note: Although the two sheets have slightly different locality data we assume they are the same collection, based on the itinerary and list of collections given for that date in Forbes's field book.

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. rockii Fosberg, Bernice P. Bishop Mus. Bull. 147: 43. 1937.
Type information:
Holotype [BISH]. J. F. Rock s.n.. October 1916. Kaua`i. Isotype(s) [BISH!].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. russi (Fosberg) Fosberg, Brittonia 8: 176. 1956. Flora of the Hawaiian Islands --Basionym
Type Information
Gouldia terminalis (Hook. & Arn.) Hillebr. f. russii Fosberg, Bernice P. Bishop Mus. Bull. 147: 49 1937.
O`ahu: Waianae Mts., Honoliuli, ridge above Kupehau, moist forest. 700 m. 30 June 1935. F.R. Fosberg 10981.
Holotype [BISH].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.
---------------------


Note: An isonym of Gouldia terminalis f. russii Fosb., Bernice P. Bishop Mus. Bull. 147: 49. 1937. (Per DHN; see comment by DNH in ARB 390, 1994).

*Verification: R.K. Shannon.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. russii Fosberg, Bernice P. Bishop Mus. Bull. 147: 49. 1937.
Type information:
Holotype [BISH]. F.R. Fosberg 10981. 30 June 1935. O`ahu: Waianae Mts., Honoliuli, ridge above Kupehau, moist forest.700m.
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. sambucina (A. Heller) Fosberg, Bernice P. Bishop Mus. Bull. 147: 36. 1937. Flora of the Hawaiian Islands --Basionym
Type Information
Gouldia sambucina A. Heller , Minnesota Bot. Stud. 1: 898 1897.
Kaua`i: Kaholuamanoa [Kaholuamanu], edge of plateau above Waimea. 3500 ft. 11-16 October 1895. A.A. Heller 2879.
Holotype [MIN]. Isotype(s) [NY [mfc!], US!].

Note: Heller did not cite a date. We have no evidence that other collections with this number were made on other dates.

=Kadua affinis
*Verification: Wagner, W.L. & R.K. Shannon. 1999. Brittonia 51:422-438.
---------------------


*Verification: R.K. Shannon.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. santalifolia Fosberg, Bernice P. Bishop Mus. Bull. 147: 51. 1937.
Type information:
Holotype [BISH]. C.N. Forbes 2310.M. 10 May 1920. Maui: Olowalu Valley. Isotype(s) [BISH!].

Note: Specific locality taken from Forbes's field notes. Fosberg apparently made in error in affixing his labels, labeling both sheets at BISH as "cotype." We have taken the fuller specimen, which more closely matches his description, to be the holotype, as have others.

=Kadua affinis
Note: Specific locality taken from Forbes's field notes. Fosberg apparently made in error in affixing his labels, labeling both sheets at BISH as "cotype." We have taken the fuller specimen, which more closely matches his description, to be the holotype, as have others.

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. sclerophylla Fosberg, Bernice P. Bishop Mus. Bull. 147: 35. 1937.
Type information:
Holotype [BISH]. C.N. Forbes 807.K. 7-18 July-August 1917. Kaua`i: head of Waimea Canyon, Waimea drainage basin, W side. Isotype(s) [BISH!, US!].

Note: Date from Forbes's field book.

=Kadua affinis
Note: Date from Forbes's field book.

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. storeyi Fosberg, Bernice P. Bishop Mus. Bull. 147: 52. 1937.
Type information:
Holotype [BISH]. H. St. John , J. Dunn and W. B. Storey 13266. 4 July 1933. Moloka`i: Wailau Valley, Kukuinui Ridge, moist woods.2400ft. Isotype(s) [BISH! [2]].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. subpilosa Fosberg, Bernice P. Bishop Mus. Bull. 147: 56. 1937.
Type information:
Holotype [BISH]. O. Degener 2570. 20 August 1927. Maui: [East Maui], Haleakala Crater, rainy region of Kaupo Gap. Isotype(s) [BISH!, GH, LCU, NY [mfc!], US?].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. suehiroae Fosberg, Bernice P. Bishop Mus. Bull. 147: 52. 1937.
Type information:
Holotype [BISH]. A. Suehiro s.n.. 3 July 1933. Moloka`i: N coast, E Molokai, Wailau Valley, top of steep bank.500ft.
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. teres Fosberg, Bernice P. Bishop Mus. Bull. 147: 36. 1937.
Type information:
Holotype [BISH]. J. F. Rock 5535. September 1909. Kaua`i: Kaholuamanu. Isotype(s) [US!].

Note: US sheet gives the collection number as 5536

=Kadua affinis
Note: US sheet gives the collection number as 5536

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. violetae Fosberg, Bull. Torrey Bot. Club 70: 392. 1943.
Type information:
Holotype [BISH]. V. O. Fosberg 49. 10 August 1936. Hawai`i: Kau, between W and E arms of 1907 flow, on dry lava. Isotype(s) [US!].
=Kadua affinis

*Verification: R.K. Shannon.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. wailauensis Fosberg, Bernice P. Bishop Mus. Bull. 147: 52. 1937.
Type information:
Holotype [BISH]. F.R. Fosberg 9625. 3 July 1933. Moloka`i: Wailau Valley, wet lower forest.150m.
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. f. waipioensis Fosberg, Bernice P. Bishop Mus. Bull. 147: 37. 1937.
Type information:
Holotype [B]. W. Hillebrand s.n.. 1851- 1871. Hawai`i: Waipio.
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. angustifolia Fosberg, Bernice P. Bishop Mus. Bull. 147: 43. 1937.
Type information:
Holotype [BISH]. H. St. John and F.R. Fosberg 12803. 28 December 1932. Moloka`i: ridge west of Punaula Valley.2000ft. Isotype(s) [BISH! [2]].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. antiqua Fosberg, Bernice P. Bishop Mus. Bull. 147: 54. 1937.
Type information:
Holotype [BISH]. J. F. Rock 8744. April 1911. Hawai`i: Kipuka Puaula. Isotype(s) [US].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. arborescens (Wawra) Fosberg, Bernice P. Bishop Mus. Bull. 147: 31. 1937. Flora of the Hawaiian Islands --Basionym
Type Information
Gouldia sandwicensis A. Gray var. arborescens Wawra, Flora 57: 276 1874.
Kaua`i: Hanalei Valley. 1869- 1870. H. Wawra 2005.
Holotype [W].
=Kadua affinis

*Verification: None.
---------------------


*Verification: R.K. Shannon.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. aspera Fosberg, Brittonia 8: 175. 1956.
Type information:
Holotype [NY]. O. Degener , T. Murashige and M. Kerr 19961. 30 August 1949. Hawai`i: Saddle Road 16 miles from Hilo, in dark, rich, rainy kipuka. Isotype(s) [BISH!, US!].

Note: Published as collection number 19161, but specimens say 19961.

=Kadua affinis
Note: Published as collection number 19161, but specimens say 19961.

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. bobeoides Fosberg, Bernice P. Bishop Mus. Bull. 147: 37. 1937.
Type information:
Holotype [BISH]. C.N. Forbes 6.L. 5 June 1913. Lana`i: Kaiholena Valley. Isotype(s) [BISH!].

Note: Collection day taken from Forbes's field notes.

=Kadua affinis
Note: Collection day taken from Forbes's field notes.

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. congesta Fosberg, Bernice P. Bishop Mus. Bull. 147: 55. 1937.
Type information:
Holotype [BISH]. C.N. Forbes 55.H. 8-14 June 1911. Hawai`i: Hualalai, Pu`uwa`awa`a. Isotype(s) [BISH!, US!].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. cordata (Wawra) Fosberg, Bernice P. Bishop Mus. Bull. 147: 44. 1937. Flora of the Hawaiian Islands --Basionym
Type Information
Gouldia sandwicensis A. Gray var. cordata Wawra, Flora 57: 277 1874.
Maui: s.l.. 1869- 1870. H. Wawra 2526.
Holotype [W].
=Kadua affinis

*Verification: None.
---------------------


*Verification: R.K. Shannon.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. coriacea (Hook. & Arn.) Fosberg, Bernice P. Bishop Mus. Bull. 147: 47. 1937. Flora of the Hawaiian Islands --Basionym
Type Information
Petesia? coriacea Hook. & Arn. , Bot. Beechey Voy. 85 1832.
O`ahu: Ko`olau Range. s.d.. G.T. Lay and A. Collie s.n..
Holotype [K].
=Kadua affinis

*Verification: None.
---------------------


*Verification: R.K. Shannon.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. crassicaulis Fosberg, Bernice P. Bishop Mus. Bull. 147: 56. 1937.
Type information:
Holotype [BISH]. O. Degener 9473. 7 July 1927. Maui: [East Maui, N slopes of Haleakala], O`opuola Stream in vicinity of Ditch Trail, in forest. Isotype(s) [BISH! [2], MASS!, MO!, NY [mfc!], US!].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. degeneri Fosberg, Bernice P. Bishop Mus. Bull. 147: 39. 1937.
Type information:
Holotype [BISH]. D.W. Garber and C.N. Forbes 166. 12 January 1920. O`ahu: Koolau Mts., right fork Wailupe Valley.
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. elongata (A. Heller) Fosberg, Bernice P. Bishop Mus. Bull. 147: 32. 1937. Flora of the Hawaiian Islands --Basionym
Type Information
Gouldia elongata A. Heller , Minnesota Bot. Stud. 1: 897 1897.
Kaua`i: [ridge] between Hanapepe and Wahiawa Rivers, [in wet boggy woods]. 2500 ft. 15 July 1895. A.A. Heller 2606.
Holotype [MIN].

Note: Collection 2606, pro parte. There is a discrepancy between the date Heller cited (25 July) and the date on the MIN sheet (15 July). The word "type" is written on the label of the MIN sheet, and we take it to be the holotype, assuming that the date cited by Heller is an error. Other collections distributed under Heller 2606 were collected at the head of the valley opposite Gay & Robinson's Hanapepe valley house, 23 July (explicitly excluded by Heller, citing them under Kadua knudsenii); between the Hanapepe and Wahiawa Rivers, 25 July (BISH! [2], NY [2], US!); and on the ridge west of the Hanapepe River, 22 August (MO!, US!), and are not types. There are two sheets at BISH that are possible isotypes, but these could not be located in August 1997.

=Kadua affinis
*Verification: Wagner, W.L. & R.K. Shannon. 1999. Brittonia 51:422-438.
---------------------


*Verification: R.K. Shannon.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. forbesii Fosberg, Bernice P. Bishop Mus. Bull. 147: 57. 1937.
Type information:
Holotype [BISH]. C.N. Forbes 947.H. 26 June 1915. Hawai`i: above Hilo, Kiipu [Puu Kipu, Keawewai]. Isotype(s) [BISH!].

Note: Check locality on map.

=Kadua affinis
Note: Check locality on map.

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. glabra Fosberg, Bernice P. Bishop Mus. Bull. 147: 36. 1937.
Type information:
Holotype [BISH]. F.R. Fosberg 10434. 10 December 1933. Hawai`i: Hilo District, Puu Oo Trail above Olaa flume, wet kipuka in 1881 flow.800m. Isotype(s) [BISH!].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. hathewayi Fosberg, Brittonia 8: 174. 1956.
Type information:
Holotype [US]. W. H. Hatheway 322. s.d.. O`ahu: Waianae Range, Mokuleia, Keawapulau Gulch, moist, shaded ravine.455m.
=Kadua affinis

*Verification: None.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. hosakai Fosberg, Bernice P. Bishop Mus. Bull. 147: 38. 1937.
Type information:
Holotype [BISH]. H. St. John , E.Y. Hashimoto and E.Y. Hosaka, J.C. Lindsay, & 11330. 26 December 1931. Hawai`i: Ka`u District; Manuka mauka, wooded kipuka.1600ft. Isotype(s) [BISH!, US!].

Note: Additional collector D.D. Mitchell.

=Kadua affinis
Note: Additional collector D.D. Mitchell.

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. kaala Fosberg, Bernice P. Bishop Mus. Bull. 147: 49. 1937.
Type information:
Holotype [BISH]. F.R. Fosberg 9098. 8 January 1933. O`ahu: Waianae Mts., Mt. Ka`ala, E slope, Waianaeuka, wet forest.1170m. Isotype(s) [US!].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. kapuaensis Fosberg, Bernice P. Bishop Mus. Bull. 147: 39. 1937.
Type information:
Holotype [BISH]. J. F. Rock s.n.. s.d.. Hawai`i: Kapua, Kona.
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. konaensis Fosberg, Bernice P. Bishop Mus. Bull. 147: 37. 1937.
Type information:
Holotype [BISH]. J. F. Rock 10031. 28 January 1912. Hawai`i: Ka`u, Manuka.

Note: Only part of this collection is type for this entity; the other part was used as the type for Gouldia terminalis f. pittosporoides Fosb. Apparently Rock's name Gouldia volcanica was never published.

=Kadua affinis
Note: Only part of this collection is type for this entity; the other part was used as the type for Gouldia terminalis f. pittosporoides Fosb. Apparently Rock's name Gouldia volcanica was never published.

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. lanai Fosberg, Bernice P. Bishop Mus. Bull. 147: 59. 1937.
Type information:
Holotype [BISH]. C.N. Forbes 80.L. 9 June 1913. Lana`i: mts near Ko`ele, up valley on middle of right hand ridge. Isotype(s) [BISH!].

Note: Collection day and specific locality taken from Forbes's field notes.

=Kadua affinis
Note: Collection day and specific locality taken from Forbes's field notes.

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. macrocarpa (Hillebr.) Fosberg, Bernice P. Bishop Mus. Bull. 147: 34. 1937. Flora of the Hawaiian Islands --Basionym
Type Information
Gouldia macrocarpa Hillebr. , Fl. Hawaiian Isl. 170 1888.
Kaua`i: s.l.. s.d.. V. Knudsen s.n..
Lectotype [B]. Lectotypified in Fosberg. Bernice P. Bishop Mus. Bull. 147: 34. 1937.

Note: Additional syntype: "O`ahu, Mt. Ka`ala!" Photo of lectotype: BISH!

=Kadua affinis
*Verification: R.K. Shannon, W.L. Wagner.
---------------------


*Verification: R.K. Shannon.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. macrothyrsa Fosberg, Bernice P. Bishop Mus. Bull. 147: 40. 1937.
Type information:
Holotype [BISH]. F.R. Fosberg 10818. 24 February 1935. O`ahu: Waianaeuka, Haleauau Valley, Puu Kaala.850m.

Note: Not in BISH type collection.

=Kadua affinis
Note: Not in BISH type collection.

*Verification: None.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. myrsinoidea Fosberg, Brittonia 8: 176. 1956.
Type information:
Holotype [NY?]. O. Degener 20966. Hawai`i: central plateau, Pu`u Keaniu.

Note: Publication indicates that type was in Degener's possession at time of publication; it was not found in the type collection or the general collection at BISH (RKS, 8/95), so presumably it is at NY.

=Kadua affinis
Note: Publication indicates that type was in Degener's possession at time of publication; it was not found in the type collection or the general collection at BISH (RKS, 8/95), so presumably it is at NY.

*Verification: None.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. osteocarpa Fosberg, Bernice P. Bishop Mus. Bull. 147: 33. 1937.
Type information:
Holotype [BISH]. H. St. John , F.R. Fosberg and & V. M. Oliveira 13943. 1 January 1934. Kaua`i: Wainiha, Wainiha Valley, moist lower forest.850ft. Isotype(s) [BISH! [2], US!].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. ovata (Wawra) Fosberg, Bernice P. Bishop Mus. Bull. 147: 50. 1937. Flora of the Hawaiian Islands --Basionym
Type Information
Gouldia sandwicensis A. Gray var. ovata Wawra, Flora 57: 278 1874.
Maui: Waihe`e Mt.. 1869- 1879. H. Wawra 1953.
Holotype [W].
=Kadua affinis

*Verification: None.
---------------------


*Verification: R.K. Shannon.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. parvifolia (Wawra) Fosberg, Bernice P. Bishop Mus. Bull. 147: 56. 1937. Flora of the Hawaiian Islands --Basionym
Type Information
Gouldia sandwicensis A. Gray var. parvifolia Wawra, Flora 57: 296 1874.
Maui: Haleakala, in forest. 1869- 1870. H. Wawra 1931.
Holotype [W].
=Kadua affinis

*Verification: None.
---------------------


*Verification: R.K. Shannon.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. pedunculata Fosberg, Bernice P. Bishop Mus. Bull. 147: 46. 1937.
Type information:
Holotype [BISH]. J. F. Rock 8363. June 1910. Hawai`i: Kohala Mts., Kohala. Isotype(s) [BISH!, US!].

Note: Fosberg cited three parts of Rock 8363, but only one as type. Others (one sheet at US, annotated as an isotype by FRF, and one at BISH) should be considered isotypes (according to DHN). Three other sheets of Rock 8363 were specifically excluded from this taxon by Fosberg, who considered them to be a hybrid between this taxon and Gouldia hillebrandii var. hawaiiensis.

=Kadua affinis
Note: Fosberg cited three parts of Rock 8363, but only one as type. Others (one sheet at US, annotated as an isotype by FRF, and one at BISH) should be considered isotypes (according to DHN). Three other sheets of Rock 8363 were specifically excluded from this taxon by Fosberg, who considered them to be a hybrid between this taxon and Gouldia hillebrandii var. hawaiiensis.

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. pseudodichotoma Fosberg, Bernice P. Bishop Mus. Bull. 147: 58. 1937.
Type information:
Holotype [BISH]. C.N. Forbes 253.L. June 1913. Lana`i: mts E end. Isotype(s) [BISH!, US!].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. pubescens Fosberg, Bernice P. Bishop Mus. Bull. 147: 57. 1937.
Type information:
Holotype [BISH]. C.N. Forbes 1898.M. 8 March 1920. Maui: [East Maui], S slope of Haleakala, Pu`u Poni. Isotype(s) [BISH!].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. pubistipula Fosberg, Brittonia 20: 291. 1968.
Type information:
Holotype [US]. O. Degener , I. Degener and W. Fleming 25310. 4 June 1959. Maui: [East Maui], Haleakala, Paliku, open forest. Isotype(s) [A, B, BISH!, G, NY [mfc!], US, W].

Note: Fleming not included as collector in publication, only on label.

=Kadua affinis
Note: Fleming not included as collector in publication, only on label.

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. purpurea Fosberg, Bernice P. Bishop Mus. Bull. 147: 46. 1937.
Type information:
Holotype [BISH]. F.R. Fosberg 10224. 3 September 1933. Hawai`i: Kohala Mts, Upper Hamakua Ditch Trail, head of Alakahi Fork of Waipio Valley, wet forest.1075m.
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. quadrangularis Fosberg, Bernice P. Bishop Mus. Bull. 147: 58. 1937.
Type information:
Holotype [BISH]. H. St. John 11378. 29 December 1931. Hawai`i: Puuwaawaa, Puu Hualalai, moist woods.5500ft. Isotype(s) [BISH! [2], US!].

Note: Other collectors: R. S. Bean, J. W. Coulter, E. Y. Hashimoto, E. Y. Hosaka, J. C. Lindsay, D. D. Mitchell.

=Kadua affinis
Note: Other collectors: R. S. Bean, J. W. Coulter, E. Y. Hashimoto, E. Y. Hosaka, J. C. Lindsay, D. D. Mitchell.

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. rotundifolia Fosberg, Bernice P. Bishop Mus. Bull. 147: 41. 1937.
Type information:
Holotype [BISH]. H. St. John and F.R. Fosberg 12911. 29 December 1932. Moloka`i: trail on ridge E of Mapulehu Valley, lower forest.1900ft. Isotype(s) [BISH! [2], US!].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. sclerotica Fosberg, Brittonia 8: 175. 1956.
Type information:
Holotype [NY]. O. Degener , T. Murashige and Kerr 20199. 28 August 1949. Hawai`i: 21 miles from Hilo, on Saddle Road, rainy kipuka. Isotype(s) [BISH!, US!].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. skottsbergii Fosberg, Bernice P. Bishop Mus. Bull. 147: 42. 1937.
Type information:
Holotype [GB]. C. Skottsberg 86. 24 September 1926. Hawai`i: S slope of Mauna Loa, Koa Camp.
=Kadua affinis

*Verification: None.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. stipulcea (Wawra) Fosberg, Bernice P. Bishop Mus. Bull. 147: 42. 1937. Flora of the Hawaiian Islands --Basionym
Type Information
Gouldia sandwicensis A. Gray var. stipulacea Wawra, Flora 57: 297 1874.
Kaua`i: plateau of Wai`ale`ale. 2440 m. 1869- 1870. H. Wawra 2176.
Holotype [W].
=Kadua affinis

*Verification: None.
---------------------


Note: Check reference for basionym, Fosberg cites it as Wawra in Hillebrand.

*Verification: R.K. Shannon.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. subcordata Fosberg, Bernice P. Bishop Mus. Bull. 147: 44. 1937.
Type information:
Holotype [BISH]. J. F. Rock 8098. 6 August 1910. Lana`i: Mahana Valley. Isotype(s) [BISH! [2], US!].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. tenuicaulis Fosberg, Bernice P. Bishop Mus. Bull. 147: 57. 1937.
Type information:
Holotype [BISH]. U. Faurie 345. March 1910. Kaua`i: Waimea.1000m.
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.

Gouldia terminalis (Hook. & Arn.) Hillebr. var. wawrana Fosberg, Bernice P. Bishop Mus. Bull. 147: 30. 1937.
Type information:
Holotype [BISH]. F.R. Fosberg 9529. 14 May 1933. O`ahu: Koolau Range, Waikane-Schofield Trail, divide between Wahiawa and Waianaeuka, wet forest.700m.
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.