Gouldia ovata (Wawra) Skottsb. , Acta Horti Gothob. 15: 465. 1944. Flora of the Hawaiian Islands --Basionym
Type Information
Gouldia sandwicensis A. Gray var. ovata Wawra, Flora 57: 278 1874.
Maui: Waihe`e Mt.. 1869- 1879. H. Wawra 1953.
Holotype [W].
=Kadua affinis

*Verification: None.
---------------------


*Verification: None.

Gouldia ovata (Wawra) Skottsb. f. russii (Fosberg) O. Deg. & I. Deg., Phytologia 7: 466. 1961. Flora of the Hawaiian Islands --Basionym
Type Information
Gouldia terminalis (Hook. & Arn.) Hillebr. f. russii Fosberg, Bernice P. Bishop Mus. Bull. 147: 49 1937.
O`ahu: Waianae Mts., Honoliuli, ridge above Kupehau, moist forest. 700 m. 30 June 1935. F.R. Fosberg 10981.
Holotype [BISH].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.
---------------------


*Verification: R.K. Shannon.

Gouldia ovata (Wawra) Skottsb. var. heterophylla (Fosberg) O. Deg. & I. Deg., Phytologia 7: 466. 1961. Flora of the Hawaiian Islands --Basionym
Type Information
Gouldia terminalis (Hook. & Arn.) Hillebr. f. heterophylla Fosberg, Bernice P. Bishop Mus. Bull. 147: 52 1937.
Moloka`i: trail on ridge E of Mapulehu Valley, wet forest. 2100 ft. 29 December 1932. H. St. John and F.R. Fosberg 13091.
Holotype [BISH]. Isotype(s) [BISH!].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.
---------------------


Note: Also published superfluously by Degener & I. Degener, 1967, Phytologia 14: 214.

*Verification: R.K. Shannon.

Gouldia ovata (Wawra) Skottsb. var. kalaupapa (Fosberg) O. Deg. & I. Deg., Phytologia 7: 466. 1961. Flora of the Hawaiian Islands --Basionym
Type Information
Gouldia terminalis (Hook. & Arn.) Hillebr. f. kalaupapa Fosberg, Bernice P. Bishop Mus. Bull. 147: 51 1937.
Moloka`i: on cliffs above Kalaupapa, Settlement Trail. June 1912. C.N. Forbes 25.Mo.
Holotype [BISH]. Isotype(s) [BISH! [3], US!].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.
---------------------


Note: Also published superfluously by Degener & I. Degener, 1967, Phytologia 14: 214.

*Verification: R.K. Shannon.

Gouldia ovata (Wawra) Skottsb. var. lydgatei (Fosberg) O. Deg. & I. Deg., Phytologia 7: 466. 1961. Flora of the Hawaiian Islands --Basionym
Type Information
Gouldia terminalis (Hook. & Arn.) Hillebr. f. lydgatei Fosberg, Bernice P. Bishop Mus. Bull. 147: 53 1937.
Maui: Hamakua. 1851- 1871. W. Hillebrand and J.M. Lydgate 126.
Holotype [BISH].

Note: The sheet containing the holotype represents a mixed collection of three different specimens. The twig in the upper left is not a Rubiaceae; the twig in the upper right is apparently from a different locality; only the twig in the lower left is the holotype. The publication indicates that there is a photo of the type at BISH, but it could not be located (RKS, 8/95). There is a photo of a sheet that Fosberg labelled "isotype," but it cannot be an isotype because it is from O`ahu, and the type was collected on Maui.

=Kadua affinis
*Verification: W.L. Wagner.
---------------------


Note: Also published superfluously by Degener & I. Degener, 1967, Phytologia 14: 214.

*Verification: R.K. Shannon.

Gouldia ovata (Wawra) Skottsb. var. makawaoensis (Fosberg) O. Deg. & I. Deg., Phytologia 7: 466. 1961. Flora of the Hawaiian Islands --Basionym
Type Information
Gouldia terminalis (Hook. & Arn.) Hillebr. f. makawaoensis Fosberg, Bernice P. Bishop Mus. Bull. 147: 50 1937.
Maui: E Maui, Haleakala, Makawao. September 1910. J. F. Rock 8531.
Holotype [BISH]. Isotype(s) [BISH!].

Note: Apparently Rock's name Gouldia haleakalensis was never published.

=Kadua affinis
*Verification: R.K. Shannon, W.L. Wagner.
---------------------


Note: Also published superfluously by Degener & I. Degener, 1967, Phytologia 14: 214.

*Verification: R.K. Shannon.

Gouldia ovata (Wawra) Skottsb. var. maunahui (Fosberg) O. Deg. & I. Deg., Phytologia 7: 466. 1961. Flora of the Hawaiian Islands --Basionym
Type Information
Gouldia terminalis (Hook. & Arn.) Hillebr. f. maunahui Fosberg, Bernice P. Bishop Mus. Bull. 147: 51 1937.
Moloka`i: Kawela, ridge between Maunahui, Kaunakakai, and Hanalilolilo. 3640 ft. 20 December 1932. H. St. John, R.J. Baker and J.W. Coulter, F.R. Fosberg, 12284.
Holotype [BISH]. Isotype(s) [BISH!].

Note: Additional collector: T. G. Yuncker.

=Kadua affinis
*Verification: R.K. Shannon, W.L. Wagner.
---------------------


Note: Also published superfluously by Degener & I. Degener, 1967, Phytologia 14: 214.

*Verification: R.K. Shannon.

Gouldia ovata (Wawra) Skottsb. var. membranacea (Fosberg) O. Deg. & I. Deg., Phytologia 7: 466. 1961. Flora of the Hawaiian Islands --Basionym
Type Information
Gouldia terminalis (Hook. & Arn.) Hillebr. f. membranacea Fosberg, Bernice P. Bishop Mus. Bull. 147: 53 1937.
Maui: Olinda, along pipeline trail, in rain forest. 16 June 1927. O. Degener 9478.
Holotype [BISH]. Isotype(s) [BISH!, NY [mfc!]].

Note: On label of BISH isotype is written (in Degener's handwriting), "Dupl. to NY, Fosberg." Check at US for isotype.

=Kadua affinis
*Verification: R.K. Shannon, W.L. Wagner.
---------------------


Note: Also published superfluously by Degener & I. Degener, 1967, Phytologia 14: 214.

*Verification: R.K. Shannon.

Gouldia ovata (Wawra) Skottsb. var. oahuensis (Fosberg) O. Deg. & I. Deg., Phytologia 7: 466. 1961. Flora of the Hawaiian Islands --Basionym
Type Information
Gouldia terminalis (Hook. & Arn.) Hillebr. f. oahuensis Fosberg, Brittonia 8: 176 1956.
O`ahu: Mokuleia, Keawapilau, forest. 1800 ft. 14 December 1950. O. Degener and W. Hatheway 21172.
Holotype [NY]. Isotype(s) [BISH!, US!].

Note: Publication gives collection number as 21722, but BISH, NY, and US sheets have 21172 on the label.

=Kadua affinis
*Verification: R.K. Shannon, W.L. Wagner.
---------------------


Note: Also published superfluously by Degener & I. Degener, 1967, Phytologia 14: 214.

*Verification: R.K. Shannon.

Gouldia ovata (Wawra) Skottsb. var. obovata (Fosberg) O. Deg. & I. Deg., Phytologia 7: 466. 1961. Flora of the Hawaiian Islands --Basionym
Type Information
Gouldia terminalis (Hook. & Arn.) Hillebr. f. obovata Fosberg, Bull. Torrey Bot. Club 70: 391 1943.
Lana'i: between Maunalaei and Hauola drainages, ridge below Puu Aalii, rather dry forest. 800 m. 30 November 1935. F.R. Fosberg 12472.
Holotype [NA]. Isotype(s) [BISH!, US!].

Note: Hauola spelled "Hauula" in publication.

=Kadua affinis
*Verification: R.K. Shannon.
---------------------


Note: Also published superfluously by Degener & I. Degener, 1967, Phytologia 14: 214.

*Verification: R.K. Shannon.

Gouldia ovata (Wawra) Skottsb. var. petiolata (Fosberg) O. Deg. & I. Deg., Phytologia 7: 466. 1961. Flora of the Hawaiian Islands --Basionym
Type Information
Gouldia terminalis (Hook. & Arn.) Hillebr. f. petiolata Fosberg, Bernice P. Bishop Mus. Bull. 147: 53 1937.
Moloka`i: Kalamaula. 5 June 1916. G.C. Munro 370.
Holotype [BISH].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.
---------------------


Note: Also published superfluously by Degener & I. Degener, 1967, Phytologia 14: 214.

*Verification: R.K. Shannon.

Gouldia ovata (Wawra) Skottsb. var. punaula (Fosberg) O. Deg. & I. Deg., Phytologia 7: 466. 1961. Flora of the Hawaiian Islands --Basionym
Type Information
Gouldia terminalis (Hook. & Arn.) Hillebr. f. punaula Fosberg, Bernice P. Bishop Mus. Bull. 147: 53 1937.
Moloka`i: ridge W of Punaula Valley, wooded ridge. 1900 ft. 28 December 1932. H. St. John and F.R. Fosberg 12797.
Holotype [BISH]. Isotype(s) [BISH! [3], US!].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.
---------------------


Note: Also published superfluously by Degener & I. Degener, 1967, Phytologia 14: 214 and 1967, Phytologia 15: 51 (the latter with spelling change to "punaulana").

*Verification: R.K. Shannon.

Gouldia ovata (Wawra) Skottsb. var. russii (Fosberg) O. Deg. & I. Deg., Phytologia 15: 51. 1967. Flora of the Hawaiian Islands --Basionym
Type Information
Gouldia terminalis (Hook. & Arn.) Hillebr. f. russi (Fosberg) Fosberg, Brittonia 8: 176 1956.


Note: An isonym of Gouldia terminalis f. russii Fosb., Bernice P. Bishop Mus. Bull. 147: 49. 1937. (Per DHN; see comment by DNH in ARB 390, 1994).

=Kadua affinis
*Verification: R.K. Shannon.
---------------------


*Verification: R.K. Shannon.

Gouldia ovata (Wawra) Skottsb. var. santalifolia (Fosberg) O. Deg. & I. Deg., Phytologia 7: 466. 1961. Flora of the Hawaiian Islands --Basionym
Type Information
Gouldia terminalis (Hook. & Arn.) Hillebr. f. santalifolia Fosberg, Bernice P. Bishop Mus. Bull. 147: 51 1937.
Maui: Olowalu Valley. 10 May 1920. C.N. Forbes 2310.M.
Holotype [BISH]. Isotype(s) [BISH!].

Note: Specific locality taken from Forbes's field notes. Fosberg apparently made in error in affixing his labels, labeling both sheets at BISH as "cotype." We have taken the fuller specimen, which more closely matches his description, to be the holotype, as have others.

=Kadua affinis
*Verification: R.K. Shannon, W.L. Wagner.
---------------------


Note: Also published superfluously by Degener & I. Degener, 1967, Phytologia 14: 214.

*Verification: R.K. Shannon.

Gouldia ovata (Wawra) Skottsb. var. storeyi (Fosberg) O. Deg. & I. Deg., Phytologia 7: 466. 1961. Flora of the Hawaiian Islands --Basionym
Type Information
Gouldia terminalis (Hook. & Arn.) Hillebr. f. storeyi Fosberg, Bernice P. Bishop Mus. Bull. 147: 52 1937.
Moloka`i: Wailau Valley, Kukuinui Ridge, moist woods. 2400 ft. 4 July 1933. H. St. John, J. Dunn and W. B. Storey 13266.
Holotype [BISH]. Isotype(s) [BISH! [2]].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.
---------------------


Note: Also published superfluously by Degener & I. Degener, 1967, Phytologia 14: 214.

*Verification: R.K. Shannon.

Gouldia ovata (Wawra) Skottsb. var. suehiroae (Fosberg) O. Deg. & I. Deg., Phytologia 7: 466. 1961. Flora of the Hawaiian Islands --Basionym
Type Information
Gouldia terminalis (Hook. & Arn.) Hillebr. f. suehiroae Fosberg, Bernice P. Bishop Mus. Bull. 147: 52 1937.
Moloka`i: N coast, E Molokai, Wailau Valley, top of steep bank. 500 ft. 3 July 1933. A. Suehiro s.n..
Holotype [BISH].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.
---------------------


Note: Also published superfluously by Degener & I. Degener, 1967, Phytologia 14: 214.

*Verification: R.K. Shannon.

Gouldia ovata (Wawra) Skottsb. var. wailauensis (Fosberg) O. Deg. & I. Deg., Phytologia 14: 214. 1967. Flora of the Hawaiian Islands --Basionym
Type Information
Gouldia terminalis (Hook. & Arn.) Hillebr. f. wailauensis Fosberg, Bernice P. Bishop Mus. Bull. 147: 52 1937.
Moloka`i: Wailau Valley, wet lower forest. 150 m. 3 July 1933. F.R. Fosberg 9625.
Holotype [BISH].
=Kadua affinis

*Verification: R.K. Shannon, W.L. Wagner.
---------------------


*Verification: R.K. Shannon.