Euphorbia celastroides Boiss. var. hathewayi Sherff, Bot. Leafl. 3:4. 1951.
Type information:
Holotype [F]. W. H. Hatheway 223. 5 September 1950. O`ahu: Mokuleia, Wai`anae Mts, east slope of Keawapilau Gulch.1400ft. Isotype(s) [BISH, NY, US!].
=Euphorbia multiformis var. multiformis

*Verification: David Lorence.

Euphorbia celastroides Boiss. , in de Candolle, Prodr. 15(2): 11. 1862.
Type information:
Lectotype [P]. J. Rémy 595. 1851- 1855. Ni`ihau. Isolectotype(s) [BISH?].

Note: Additional syntype: Remy 595, Kaua`i. Sherff made the Kaua`i collection the type of E. celastroides var. humbertii. Although Boissier did not cite numbers for the Remy collections, it is presumed that they are both number 595. The BISH sheet has neither locality nor number on it, so it cannot be determined whether it is an isolectotype of E. celastroides or an isotype of E. celastroides var. humbertii, if either (RKS, 8/95). Was this lectotypified by Sherff, 1937, Annals of the Missouri Botanical Garden? WLW sez, "Need to see material." Koutnik cites Remy 595 Ni`ihau, but does the number 595 really go with both the Ni`ihau and the Kaua`i collection (Koutnik is the only one who says 595 for the Ni`ihau sheet). Is the publication of var. humbertii (Bot. Gaz. 97: 596. 1936.) using syntype material of var. celastroides 1) effective lectotypification of var. celastroides or 2) invalid publication of var. humbertii?? (11/95). Koutnik, D.L. 1987. Allertonia 4:331-388

=Euphorbia celastroides var. celastroides
Note: Additional syntype: Remy 595, Kaua`i. Sherff made the Kaua`i collection the type of E. celastroides var. humbertii. Although Boissier did not cite numbers for the Remy collections, it is presumed that they are both number 595. The BISH sheet has neither locality nor number on it, so it cannot be determined whether it is an isolectotype of E. celastroides or an isotype of E. celastroides var. humbertii, if either (RKS, 8/95). Was this lectotypified by Sherff, 1937, Annals of the Missouri Botanical Garden? WLW sez, "Need to see material." Koutnik cites Remy 595 Ni`ihau, but does the number 595 really go with both the Ni`ihau and the Kaua`i collection (Koutnik is the only one who says 595 for the Ni`ihau sheet). Is the publication of var. humbertii (Bot. Gaz. 97: 596. 1936.) using syntype material of var. celastroides 1) effective lectotypification of var. celastroides or 2) invalid publication of var. humbertii?? (11/95). Koutnik, D.L. 1987. Allertonia 4:331-388

*Verification: None.

Euphorbia celastroides Boiss. f. kahanana Sherff, Bot. Gaz. (Crawfordsville) 97: 596. 1936.
Type information:
Holotype [BISH]. O. Degener and N. Krauss 8036. 26 February 1929. O`ahu: W slope of Kahana Valley, sunny ridge. Isotype(s) [F, NY [mfc!]].

Note: Discrepancies between BISH sheet, Sherff publication, and Koutnik as to where the holotype is. BISH sheet is considered holotype (ex herb. Degener) even though it is labeled "isotype." See Sherff's later citation in Ann. Missouri Bot. Gard. 25: 31 (1938) ("type, Degener; cotypes, Field, N. Y."). R.K. Shannon

=Euphorbia celastroides var. amplectens
Note: Discrepancies between BISH sheet, Sherff publication, and Koutnik as to where the holotype is. BISH sheet is considered holotype (ex herb. Degener) even though it is labeled "isotype." See Sherff's later citation in Ann. Missouri Bot. Gard. 25: 31 (1938) ("type, Degener; cotypes, Field, N. Y."). R.K. Shannon

*Verification: David Lorence.

Euphorbia celastroides Boiss. var. amplectens Sherff, Bot. Gaz. (Crawfordsville) 97: 598. 1936.
Type information:
Holotype [F]. C.N. Forbes 355.Mo. August 1912. Moloka`i: Kalua`uha Valley. Isotype(s) [BISH! [3], F, K, MIN, MO!, US!].

Note: Koutnik, D.L. 1987. Allertonia 4:331-388

=Euphorbia celastroides var. amplectens
Note: Koutnik, D.L. 1987. Allertonia 4:331-388

*Verification: David Lorence.

Euphorbia celastroides Boiss. var. arenisaxosa H. St. John, Phytologia 63: 466. 1987.
Type information:
Holotype [BISH]. H. St. John , G. Coltam and C. F. Hudson, I. E. Lane 23486. 24 December 1948. Moloka`i: Kalani, Kaluakoi, sand dunes.200ft. Isotype(s) [BISH! [2], US!].

Note: Additional collector: J. H. Maysilles.

=Euphorbia celastroides var. amplectens
Note: Additional collector: J. H. Maysilles.

*Verification: David Lorence.

Euphorbia celastroides Boiss. var. halawana Sherff, Bot. Gaz. (Crawfordsville) 97: 596. 1936.
Type information:
Holotype [BISH]. J. F. Rock 14041. April 1918. Moloka`i: E end, near Halawa. Isotype(s) [BISH! [3], A].

Note: Koutnik, D.L. 1987. Allertonia 4:331-388

=Euphorbia celastroides var. amplectens
Note: Koutnik, D.L. 1987. Allertonia 4:331-388

*Verification: David Lorence.

Euphorbia celastroides Boiss. var. hanapepensis Sherff, Bot. Gaz. (Crawfordsville) 97: 597. 1936.
Type information:
Holotype [MO]. A.A. Heller 2429. 2-8 July 1895. Kaua`i: along the Hanapepe River near the falls.700ft. Isotype(s) [US!].

Note: Collection 2429, pro parte. Other collections distributed under Heller 2429 were made 24-26 June (A, GH, BISH! [3], MIN!, NY [mfc!]), and are not primary type material. Sherff clearly distinguished between specimens of Heller 2429 collected 24-26 June and those collected 2-8 July. He cited only the US sheet as an isotype. Possible isotypes: CU, MSC. The paratypes cited by Sherff (F [2], GH, K, NY) probably all represent the same collection, although the locality on the K sheet is given as "Hanapepe and Wahiawa watershed," and the date is 24 June. Koutnik (1987) erroneously considered specimens at BISH, F, GH, K, and MIN to be isotypes. Wagner, W.L. & R.K. Shannon. 1999. Brittonia 51:422-438

=Euphorbia celastroides var. hanapepensis
Note: Collection 2429, pro parte. Other collections distributed under Heller 2429 were made 24-26 June (A, GH, BISH! [3], MIN!, NY [mfc!]), and are not primary type material. Sherff clearly distinguished between specimens of Heller 2429 collected 24-26 June and those collected 2-8 July. He cited only the US sheet as an isotype. Possible isotypes: CU, MSC. The paratypes cited by Sherff (F [2], GH, K, NY) probably all represent the same collection, although the locality on the K sheet is given as "Hanapepe and Wahiawa watershed," and the date is 24 June. Koutnik (1987) erroneously considered specimens at BISH, F, GH, K, and MIN to be isotypes. Wagner, W.L. & R.K. Shannon. 1999. Brittonia 51:422-438

*Verification: David Lorence.

Euphorbia celastroides Boiss. var. haupuana Sherff, Bot. Gaz. (Crawfordsville) 97: 594. 1936.
Type information:
Holotype [BISH]. J. F. Rock 2444. 18 March 1909. Kaua`i: Haupu Lihu`e. Isotype(s) [GH].

Note: Koutnik, D.L. 1987. Allertonia 4:331-388

=Euphorbia celastroides var. celastroides
Note: Koutnik, D.L. 1987. Allertonia 4:331-388

*Verification: David Lorence.

Euphorbia celastroides Boiss. var. humbertii Sherff, Bot. Gaz. (Crawfordsville) 97: 596. 1936.
Type information:
Holotype [GH]. J. Rémy 595. 1851- 1855. Kaua`i. Isotype(s) [BISH?, GH, P [2 sheets]].

Note: Additional syntype: Remy 595, Ni`ihau. Koutnik made the Ni`ihau collection the lectotype of E. celastroides. Although Boissier did not cite numbers for the Remy collections, it is presumed that they are both number 595. The BISH sheet has neither locality nor number on it, so it cannot be determined whether it is an isolectotype of E. celastroides or an isotype of E. celastroides var. humbertii, if either (RKS, 8/95). Koutnik, D.L. 1987. Allertonia 4:331-388

=Euphorbia celastroides var. celastroides
Note: Additional syntype: Remy 595, Ni`ihau. Koutnik made the Ni`ihau collection the lectotype of E. celastroides. Although Boissier did not cite numbers for the Remy collections, it is presumed that they are both number 595. The BISH sheet has neither locality nor number on it, so it cannot be determined whether it is an isolectotype of E. celastroides or an isotype of E. celastroides var. humbertii, if either (RKS, 8/95). Koutnik, D.L. 1987. Allertonia 4:331-388

*Verification: David Lorence.

Euphorbia celastroides Boiss. var. ingrata O. Deg. & Sherff, in Sherff, Bot. Leafl. 9: 7. 1954.
Type information:
Holotype [F]. O. Degener 21967. 28 February 1952. Maui: West Maui, Kahakuloa, grassy windswept promontory. Isotype(s) [BISH!, K, NY [mfc!]].

Note: Koutnik, D.L. 1987. Allertonia 4:331-388

=Euphorbia celastroides var. amplectens
Note: Koutnik, D.L. 1987. Allertonia 4:331-388

*Verification: David Lorence.

Euphorbia celastroides Boiss. var. kaenana Sherff, Bot. Gaz. (Crawfordsville) 97: 594. 1936.
Type information:
Holotype [BISH]. V. MacCaughey s.n.. 28 March 1915. O`ahu: Kaena uplands.

Note: Koutnik, D.L. 1987. Allertonia 4:331-388

=Euphorbia celastroides var. kaenana
Note: Koutnik, D.L. 1987. Allertonia 4:331-388

*Verification: David Lorence.

Euphorbia celastroides Boiss. var. kealiana Sherff, Bot. Gaz. (Crawfordsville) 97: 595. 1936.
Type information:
Holotype [BM]. U. Faurie 477. Kaua`i: Ke`alia, in rupibus littoris. Isotype(s) [A].

Note: Koutnik, D.L. 1987. Allertonia 4:331-388

=Euphorbia celastroides var. celastroides
Note: Koutnik, D.L. 1987. Allertonia 4:331-388

*Verification: David Lorence.

Euphorbia celastroides Boiss. var. kohalana O. Deg. & Sherff, in Sherff, Bot. Gaz. (Crawfordsville) 97: 595. 1936.
Type information:
Holotype [G]. O. Degener 8037. 22 March 1930. Hawai`i: Kohala. Isotype(s) [BISH! [3], CU, F, LCU, MASS!, MICH, MO!, NY [mfc!]].

Note: Koutnik, D.L. 1987. Allertonia 4:331-388

=Euphorbia celastroides var. amplectens
Note: Koutnik, D.L. 1987. Allertonia 4:331-388

*Verification: David Lorence.

Euphorbia celastroides Boiss. var. laehiensis O. Deg., I. Deg. & Sherff, in Sherff, Occas. Pap. Bernice P. Bishop Mus. 23(7): 122. 1964.
Type information:
Holotype [F]. O. Degener and I. Degener 28760. 26 January 1964. Lana`i: Laehi, only on calcareous, rounded slope.100ft. Isotype(s) [A, B, BISH!, CU, E, F, G, K, L MASS!, MICH, MIN, NY [mfc!], MO!, US!].

Note: Koutnik (1985) erroneously published type as Degener & Degener 28375 (RKS, 8/95). Label on MO sheet indicates that Degener & Degener made the combination in Chamaesyce--check lit. to confirm that it was not published by them. Koutnik, D.L. 1985. Brittonia 37:397-399

=Euphorbia celastroides var. laehiensis
Note: Koutnik (1985) erroneously published type as Degener & Degener 28375 (RKS, 8/95). Label on MO sheet indicates that Degener & Degener made the combination in Chamaesyce--check lit. to confirm that it was not published by them. Koutnik, D.L. 1985. Brittonia 37:397-399

*Verification: David Lorence.

Euphorbia celastroides Boiss. var. limahuliensis H. St. John, Phytologia 64: 487. 1988.
Type information:
Holotype [BISH]. S. Perlman 9. 12 April 1978. Kaua`i: Limahuli Valley, SW end, on exposed slopes.1400 to 1700ft. Isotype(s) [BISH! [5]].

Note: Why so many sheets at BISH? Any dups. anywhere else? R.K. Shannon, D.R. Herbst

=Euphorbia celastroides var. hanapepensis
Note: Why so many sheets at BISH? Any dups. anywhere else? R.K. Shannon, D.R. Herbst

*Verification: David Lorence.

Euphorbia celastroides Boiss. var. lorifolia (A. Gray) Sherff, Bot. Gaz. (Crawfordsville) 97: 597. 1936. Flora of the Hawaiian Islands --Basionym
Type Information
Euphorbia multiformis Hook. & Arn. var. lorifolia A. Gray, in Mann, Proc. Amer. Acad. Arts 7: 202 1867.
Maui: sandy isthmus. 1864- 1865. H. Mann and W.T. Brigham 389.
Isotype(s)
=Euphorbia celastroides var. lorifolia

*Verification: David Lorence.
---------------------


Note: Note also, this variety publ. ex Skottsb., Acta Horti Gothob. 10: 122. 1936. [D.M. Mix]

*Verification: d.

Euphorbia celastroides Boiss. var. mauiensis Sherff, Bot. Gaz. (Crawfordsville) 97: 601. 1936.
Type information:
Holotype [US]. H. Mann and W.T. Brigham 389. 1864- 1865. Maui: sandy isthmus. Isotype(s) [BISH, CU, F, GH, MO!].

Note: Nom. illeg. This collection was used as the type for Euphorbia multiformis var. lorifolia by Gray. [W.L. Wagner, D.M. Mix]

=Euphorbia celastroides var. lorifolia
Note: Nom. illeg. This collection was used as the type for Euphorbia multiformis var. lorifolia by Gray. [W.L. Wagner, D.M. Mix]

*Verification: David Lorence.

Euphorbia celastroides Boiss. var. moomomiana Sherff, Bot. Gaz. (Crawfordsville) 97: 593. 1936.
Type information:
Holotype [BISH]. J. F. Rock 14014. April 1918. Moloka`i: Mo`omomi sands. Isotype(s) [BISH! [3], GH, K, US!].

Note: Rock's name Euphorbia munroi was apparently never published. Koutnik, D.L. 1987. Allertonia 4:331-388

=Euphorbia celastroides var. stokesii
Note: Rock's name Euphorbia munroi was apparently never published. Koutnik, D.L. 1987. Allertonia 4:331-388

*Verification: David Lorence.

Euphorbia celastroides Boiss. var. nelsonii H. St. John, Pacific Sci. 30: 15. 1976.
Type information:
Holotype [BM]. A. Menzies and D. Nelson s.n.. 1792- 1794.
=Euphorbia celastroides var. hanapepensis

*Verification: David Lorence.

Euphorbia celastroides Boiss. var. nematopoda Sherff, Bot. Gaz. (Crawfordsville) 97: 597. 1936.
Type information:
Holotype [BISH]. C.N. Forbes 726.K. 1 November 1916. Kaua`i: Ha`upu Range, left hand side of Kipu Kai Gap.

Note: Koutnik, D.L. 1987. Allertonia 4:331-388

=Euphorbia celastroides var. celastroides
Note: Koutnik, D.L. 1987. Allertonia 4:331-388

*Verification: d.

Euphorbia celastroides Boiss. var. niuensis Sherff, Bot. Gaz. (Crawfordsville) 97: 598. 1936.
Type information:
Holotype [B]. W. Hillebrand s.n.. O`ahu: Niu.

Note: Holotype presumably destroyed; photo F.

=Euphorbia celastroides var. kaenana
Note: Holotype presumably destroyed; photo F.

*Verification: David Lorence.

Euphorbia celastroides Boiss. var. odonatoides O. Deg. & Sherff, in Sherff, Amer. J. Bot. 38: 55. 1951.
Type information:
Holotype [F]. O. Degener and T. Murashige 19779. 4 August 1949. Lana`i: Waiakeakua, in decadent forest.2500ft. Isotype(s) [BISH!, BM, CU, DS, F [photo US!], GH, K, M, MIN, MO!, NY [4, mfc!], US! [2]].

Note: Additional herbaria: CM, G-DEL, LA, P, PH. There is a photo of one of the NY sheets at F. Koutnik, D.L. 1987. Allertonia 4:331-388

=Euphorbia celastroides var. lorifolia
Note: Additional herbaria: CM, G-DEL, LA, P, PH. There is a photo of one of the NY sheets at F. Koutnik, D.L. 1987. Allertonia 4:331-388

*Verification: David Lorence.

Euphorbia celastroides Boiss. var. pseudoniuensis O. Deg. & Sherff, in Sherff, Amer. J. Bot. 43: 475. 1956.
Type information:
Holotype [F]. O. Degener , I. Degener and H. W. Hansen, & G. Cliff 23989. 23 September 1955. Kaua`i: Koke`e, Nu`alolo Trail, open ridge. Isotype(s) [BISH! [2], F, K, NY [2, mfc!]].

Note: Koutnik, D.L. 1987. Allertonia 4:331-388

=Euphorbia atrococca
Note: Koutnik, D.L. 1987. Allertonia 4:331-388

*Verification: David Lorence.

Euphorbia celastroides Boiss. var. saxicola O. Deg. & Sherff, in Sherff, Bot. Gaz. (Crawfordsville) 97:596. 1936.
Type information:
Holotype [G]. O. Degener 8088. 9 August 1926. Hawai`i: Kohala, along rocky shore. Isotype(s) [BISH! [2], CU, F, LCU].
=Euphorbia celastroides var. amplectans

*Verification: Koutnik, D.L. 1987. Allertonia 4:331-388.

Euphorbia celastroides Boiss. var. stokesii (C. N. Forbes) Sherff, Bot. Gaz. (Crawfordsville) 97: 593. 1936. Flora of the Hawaiian Islands --Basionym
Type Information

*Verification: David Lorence.

Euphorbia celastroides Boiss. var. tomentella (Boiss.) Oudejans, Phytologia 67: 44. 1989. Flora of the Hawaiian Islands --Basionym
Type Information
Euphorbia multiformis Hook. & Arn. var. tomentella Boiss., in de Candolle, Prodr. 15(2): 12 1862.
O`ahu. 1851- 1855. J. Rémy 594.
Holotype [P].
=Euphorbia celastroides var. tomentella

*Verification: David Lorence.
---------------------


*Verification: David Lorence.

Euphorbia celastroides Boiss. var. waikoluensis Sherff, Bot. Gaz. (Crawfordsville) 97: 594. 1936.
Type information:
Holotype [F]. J. F. Rock 6191. 26 March 1910. Moloka`i: Kalaupapa and Waikolu along the shore. Isotype(s) [BISH! [3], F, GH].

Note: Koutnik, D.L. 1987. Allertonia 4:331-388

=Euphorbia celastroides var. amplectens
Note: Koutnik, D.L. 1987. Allertonia 4:331-388

*Verification: David Lorence.