| General Information |
| Distribution | In the Hawaiian Islands,
endemic to
Kaua`i, O`ahu, Moloka`i, Lana`i, Maui.
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Habit
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Trees, small or rarely up to 20 m tall; branches yellowish to dark brown.
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Leaves
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Leaves coriaceous, narrowly oblanceolate to obovate, 5-13 cm long, 1.5-6 cm wide, lateral veins 7-12 pairs, marginal ribs often developed, lower surface usually glabrous except for the domatia, domatia usually very conspicuous, up to 3 mm long (particularly in the Kaua`i and some O`ahu representatives), and usually filled with whitish hairs, apex acute to obtuse or rounded, usually with an obtuse, abrupt point, base usually acuminate or attenuate, petioles 0.5-3.3 cm long, stipules broadly ovate to obovate, up to 10 mm long, often as wide as long or wider.
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Flowers
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Flowers functionally unisexual, 4-6-merous, inflorescences with 1 main axis and 2-3 nodes of usually verticillate branching, usually not pendent, branches 2-22 mm long, those of the upper node or nodes much smaller than those of the lowermost node, peduncles robust, 1.2-6.2 cm long; calyx usually thick and fleshy, 1.5-4 mm long, somewhat or not at all dilated toward apex, glabrous, truncate or with obsolete lobes; corolla tube 1-2.5 mm long, barely or not at all expanded toward apex, throat glabrous or pubescent, the lobes usually thick and fleshy, broadly ovate, shorter than tube, reflexed; filaments of staminate flowers up to 1 mm long at anthesis; anthers attached at base, ovate-oblong, 1-1.2 mm long; ovary usually turbinate, 1.2-2.5 mm long; style 0.5-0.8 mm long; stigmas 2, thick, about as long as style or slightly smaller, glabrous or papillose.
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Fruit
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Fruit pyriform to oblong-globose, 10-12 mm long, pyrenes hemispherical in cross section, usually with a T-shaped invagination of the seed coat on the ventral surface.
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