Flora of the Hawaiian Islands
Monocotyledon Literature for Cyperus laevigatus
Wagner et al., 1990, 1999.
   Cyperaceae -- The Sedge Family Bibliography
      Cyperus laevigatus

Common name(s): umbrella segde, ehu`awa, makaloa, makoloa
General Information
DistributionWidespread in warm temperate and subtropical regions.In the Hawaiian Islands, indigenous to Laysan, Ni`ihau, O`ahu, Moloka`i, Maui, Hawai`i.

Cyperaceae - Cyperus laevigatus















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Habit
Perennials, rhizomes creeping, short or elongate, clothed with brown scales; culms arranged in a single row along the rhizome, widely or closely spaced, smooth, moderately stout, trigonous, 10-45 cm tall.
Leaves
Leaves reduced to bladeless sheaths; sheaths 2-3, brown or dark brown, obliquely truncate at orifice, the uppermost sheath with a short blade, the blade green, glaucous, subulate, 3-30 mm long, convolute, apex mucronate.
Flowers
Inflorescences anthelate, capitate with 1-12 sessile spikelets; involucral bracts 2, the longer one culm-like, erect, 3-8 cm long, much longer than the inflorescence, the shorter one spreading; spikelets 12-24-flowered, pale yellowish brown, lanceolate-oblong, slightly swollen, 7-12 mm long, ca. 2 mm wide, apex acute, rachilla tetragonal, not winged; glumes pale yellowish brown, herbaceous, closely spaced, broadly ovate, 2-2.3 mm long, occasionally with rusty brown striations, 3-4-nerved on the dorsally depressed keel, apex obtuse or mucronulate; style short; stigmas 2.
Fruit
Achenes grayish brown, obovate to ovate, dorsiventrally lenticular or convexo-concave, ca. 1.5 mm long, puncticulate, apex obtuse.
Chromosomes
2n = 72, 86, 88