Flora of the Hawaiian Islands
Monocotyledon Literature for Bothriochloa pertusa
Wagner et al., 1990, 1999; Lorence & Wagner, 2019.
   Poaceae -- The Grass Family Bibliography
      Bothriochloa pertusa

Common name(s): beardgrass, pitted beardgrass
General Information
DistributionNative to the Paleotropics, now naturalized in Neotropical areas.In the Hawaiian Islands, naturalized on Midway, Ni`ihau, Kaua`i, O`ahu, Moloka`i, Lana`i, Maui, Kaho`olawe, Hawai`i.

Sprawling perennial, rhizomes contracted; culms 30‒100 cm tall, hollow, freely branching, glabrous except upwardly bearded at nodes, rooting at proximal nodes.
Leaves with keeled sheath, hirsute; ligule a ciliate membrane, 0.7‒1.2 mm long; blade 3‒4 mm wide, with scattered, elongate, papillose based hairs along margin and above ligule.
Inflorescences terminal, often purplish, flabellate, 3‒5 cm long, peduncle exserted up to 12 cm from the subglabrous, bladeless distal sheath, glabrous or slightly bearded at apex, consisting of several ascending racemes, racemosely arranged along a short axis, pedicel of pedicellate spikelet ca. 3 mm long, densely villous, flattened. Pedicellate spikelet 3‒4 mm long, sterile or sometimes staminate, often purple tinged, scaberulous; sessile spikelet ca. 5 mm long; first glume subcoriaceous, with a conspicuous pit in the middle, villous on proximal ½, scabrous on distal ½, attenuate, often bidentate at apex, second glume ca. 5 mm long, submembranous, keeled, minutely puberulent at apex, otherwise glabrous, apiculate; first (sterile) lemma 2.5‒2.7 mm long, hyaline, ciliate on distal margin, fertile lemma reduced to a slender, geniculate, reddish brown awn 15‒20 mm long; palea absent.
Caryopses ovoid to ellipsoid, 1.5‒2 mm long.
2n = 36, 40, 60.
Nancy Khan