Flora of the Hawaiian Islands
Pteridophyte
   Cystopteidaceae
      Cystopteris -- The bladder fern genus
General Information
DistributionA genus of about 25-30 species widespread in temperate zones and at higher elevations in the Tropics.
Habit
Epipetric or Terrestrial, small to medium-sized; rhizomes short-creeping to decumbent.
Leaves
Stipes slender, grooved, straw-colored to medium brown, mostly glabrous, bases with occasional membranous scales; cross-sections at bases reveal ends of 2 ribbon-shaped vascular bundles (appearing as linear and parallel lines); blade 2-pinnate to 2-pinnate-pinnatifid, herbaceous to somewhat coriaceous, glabrous or slightly scaly; veins free.
Sori
Sori round to cup-shaped; indusia hoodlike, attached at bases, free at margins, opening outward, partially overlapping sori.
Notes
Greek kystos, bladder or sac, + pteris, fern, alluding to the indusium shape.
Contributor
Nancy Khan