Leaves
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Fronds distant (to 10 cm apart), monomorphic, pinnate-pinnatifid, erect or arching; stipe blackish at base, stramineous distally, adaxially grooved, scales sparse, ovate-lanceolate, brown, setulose on margins and glabrous on surfaces; blade chartaceous to subcoriaceous (often somewhat leathery), drying greenish or reddish, pinnate-pinnatifid, with proximal pinnae not reduced or lowermost pair only slightly so, lacking auricles at acroscopic base, apex usually short, pinnatifid to subconform and pinna-like but usually much shorter than longest lateral pinnae; rachis generally hairy to glabrous or glabrescent, hairs if present 0.1–1.0 mm long, not scaly, lacking proliferous buds in axils of pinnae; pinnae 10–30 pairs, linear-lanceolate, short-stalked (1–4 mm), usually alternate or becoming alternate distally, lacking auricles, bases rounded to truncate, adaxially with a groove that is not continuous with the rachis groove, shallowly lobed ca.
1⁄4–1⁄2(-5/6) their width; veins simple, 9–18(–30) pairs per segment, usually prominent on both sides, unbranched, basal pairs from adjacent segments united at an obtuse angle below the sinus, and forming an excurrent vein (usually 2–4 mm long) running to the sinus, next pair more oblique and meeting margin at or just above the sinus, free vein ends reaching segment margins; aerophores inconspicuous or absent at pinna bases, not swollen; indument abaxially of stipe, rachis, costa, veins, and often laminar tissue between veins lacking or of hyaline acicular, spreading hairs, these 0.1–0.4 mm, blades lacking capitate hairs, often with orangish to reddish, sessile, spherical glands on costules, veins, and sometimes on laminar tissue between veins, also with scattered ovate, flat, pale brown scales on costae and sometimes on costules and veins; indument adaxially of generally short (ca. 0.2 mm) hyaline, unicellular setae on stipe, rachis, and costa, hairs lacking on veins and tissue between veins; pustules absent on laminar tissue on both sides.
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