Flora of the Hawaiian Islands
Dicotyledon
   Asteraceae -- The Sunflower Family
      Bidens -- The beggartick, ko`oko`olau, ko`olau, Spanish needle genus
General Information
DistributionA taxonomically difficult genus of about 340 species. The largest numbers are native to the Americas, Africa, and Polynesia, with a few in Europe and Asia. Several weedy species have become naturalized worldwide.
Habit
Annual or perennial herbs or shrubs, 0.1‒4 m tall.
Leaves
Leaves simple and serrate or pinnately to tripinnately compound, opposite, pubescent or glabrous, petiolate.
Flowers
Heads solitary on long peduncle or in simple to compound cymes terminating the main stem and lateral branches or only on lateral branches, radiate or discoid; involucral bracts in 2 series, outer ones green and herbaceous, inner ones membranous or chaffy, rarely green and connate; receptacle chaffy; bracts usually linear, flat or nearly so; ray florets absent or 3‒19 per head, sterile, rays yellow or white, rarely orange or red; disk florets perfect or pistillate (and then plants gynodioecious), corollas yellow or orange, (4)5-lobed; tubular or urceolate nectary present, surrounding base of style; anthers sagittate with small, smooth auricles or entire; style branches tipped with short, acute or longer subulate appendages; pappus of 0‒4(‒8) barbed or glabrous awns.
Fruit
Achenes obovate-oblong, cuneate, or linear, obcompressed or 3‒4-angled, straight, curved, twisted, or coiled, occasionally with lateral wings, setose or glabrous.
Notes
See Wagner et al. (2014) for a revision of the endemic Marquesas Bidens. The name is derived from the Latin bi, two, and dens, teeth, in refrence to the usually paired pappus awns.
Contributor
Nancy Khan